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The tiny remote islands of Galápagos have played a central role in the study of evolution. Not surprisingly, they have also been important to theory and practice in biodiversity conservation. The fascinating adaptations of organisms to the unusual, isolated ecosystems of the archipelago have left them particularly vulnerable to perturbations and introductions from the outside. Drawing on lessons learned from Darwin's time to the present, this seminar explores evolution, conservation, and their connection among the habitats and organisms of Galápagos. Using case-study material on tortoises, iguanas, finches, Scalesia plants, penguins, cormorants and more, we will explore current theory and debate about adaptation, speciation, adaptive radiation, sexual selection, and other topics in evolution. Similarly, we will explore the special challenges Galápagos poses today for conservation, owing to both its unusual biota and to the increasing impact of human activity in the archipelago. This course includes an intensive ten-day expedition to Galápagos at no additional cost to students. The goal of the expedition is both to observe firsthand many of the evolutionary adaptations and conservation dilemmas that we have read about, and to look for new examples and potential solutions. A chartered ship named MV La Pinta will serve as our floating classroom, dormitory, and dining hall as we work our way around the archipelago to visit seven different islands (Darwin visited four). For this portion of the class, undergraduates will be joined by a small group of Stanford alumni and friends in a format called a Stanford "Field Seminar." Because our class time on campus is limited to one week before travel, students will be required to complete all course readings over the summer. The course emphasizes student contributions and presentations. Students will be asked to lead class discussions and to carry out a thorough literature review of some aspect of the evolution and/or conservation of one or more Galápagos species. The final assignment for the seminar is to complete an eight- to ten-page paper about that review and to present its main findings in a joint seminar of undergrads and alumni as we travel in Galápagos. The class is also an experiment in intergenerational learning. For a few days on campus, and for all travels, we will be joined by 20+ Stanford alumni and friends who will share in our educational activities. For many alums, this class will be their first experience back in the Quad classrooms since their days as students, and much has changed at Stanford in the meantime. The class requires an attitude of mutual respect and a willingness to learn from each other on the part of each person in both groups, students and alumni. We will ask you to demonstrate that you respect each other's perspectives by listening and sharing what you are learning from each other; and by communicating about...(see complete course description at soco.stanford.edu.)